Leave the springs off and try it. I don't know why a "bolt on" QR would even use them(bolt on defeats the whole quick release idea). I bet the spring is binding between the end of the axle and the inside of the QR nuts.
When I say outer edge of the drop out: What I am saying is put the wheel in the frame with out the QR in place. If the ends of the axle are very close to the outside of the frame(dropouts) then you might have an interference issue, especially if you use the springs. From what you have describes I am betting this is it. I have had this same problem when putting newer wheels on some older frames. The spring just took up to much room. Easier to remove it than cut the axle down.
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I am in the woods and I have gone crazy.