Join Date: Jun 2007
Location: Western Wisconsin, USA
Bikes: Catrike Trail (fixed frame) tadpole trike recumbent
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Compared to the supine styles that have evolved with rider on back, prone has two primary disadvantages: weight is borne on the chest which interferes with breathing, and to see forward the rider must hold their neck back at a considerable angle.
I recall pics of one-off speed experiments, but haven't heard of anyone riding one regularly. Putting the feet back by the rear wheel can save drivetrain weight. One experimenter rode on their back, head first, navigating with either a mirror or digicam/monitor.
There may be some out there doing it prone, but I doubt there's any volume to the numbers.