RDB

I'm getting new tires and the choice is between these two. Both are durable and offer relatively good puncture protection. I'll be getting the 25mm width.

Which one**********????
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Price is not a concern.

Retro Grouch 

I've used Gatorskins quite a bit in the past and been happy with them. If I was still using 700c tires today they'd probably be Gatorskins.

If I was completely honest with myself, I think that a big factor is that I liked the name. As a bonus, the majority of folks that I ride with use Gatorskins so, if I happen to puncture on a group ride, I won't have to endure a lecture about my sucky tire choice while everybody else is standing around waiting for me to fix my flat.

Cyclosaurus

Depends on what you are looking for. There is usually a tradeoff—faster rolling tires come at the cost of durability. The GP 4 Season is meant to be the middle ground between Continental's Grand Prix and Gatorskin tires. Faster and more supple than the Gatorskin, with more durability and flat protection than the Grand Prix. I use GP 4 Season 28s on my CX bike because it is a nice happy medium between the two.

Drew Eckhardt 

Gatorskin or GP4000. Similar lifetime and flat resistance if you don't damage the sidewalls.

The Gatorskin has side-wall protection and rides OK for such a flat resistant tire.

The GP4000S does not and is surprisingly durable and flat resistant for a fast nice riding tire.

The GP 4 Season lasts about half as long as either and doesn't ride appreciably better than the Gatorskin.

Frreed

I like the Gatorskins. I did an 8 day 500+ mile ride across Kansas in June. No flats at all over the course of the ride. The roads were decent chip seal but there was plenty of glass and other junk on the surface. We had a day where it rained hard for about 2 hours and they handled that quite well. The peace of mind they give me is worth any "extra" rolling resistance.

Squashednuts

I see you've narrowed it down to two choices but have you thought of Maxxis Refuse, they come in 25mm are extremely hard wearing & I find them better in the wet than Gatorskins.

shelbyfv

Between the two, I prefer the Gators. I found the 4 Season the be just as harsh riding but less durable. If you are open to another suggestion, the new GP GT rides nearly as well as the 4000sII but is more durable.

Stucky

Gatorskins. Once you get 'em, you never even have to give a thought to your tires, or worry about flats.

Cyclosaurus

Funny...there's this recent thread full of hate for Gatorskins...I guess they stir deep passions one way or another

fietsbob

'gatorskin' mesh reinforces the sidewalls .

andr0id

LOL, that's a selling point I never thought of.

WarDonkey

Don't know about the Gatorskins, but I use the GP 4 seasons. 1000 miles and going strong. I ride mostly country roads with occasional gravel road thrown in. Rode in the rain a couple of times and they seem to hold the road well.

Elvo

Michelin Pro 4 Endurance

bowzette

I like Gatorskins on the rear wheel for flat protection and very little tire wear from trainer use. I do think they ride rather "hard" which is why I usually don't use them on the front wheel.

MarkND

I've had good luck with Gatorskins. I'm on my second set and I've never gotten a flat while they have been on my bike. Before I switched to them I would get flat tires all the time. Not getting flat tires is kind of important to me, nothing is more of a ride killer than a flat tire. I'm not riding in the TDF with support vehicles never far away, so I'll stick with my Gatorskins.

Mark

Jarrett2

I ride on really bad roads daily. I like the Gatorskins, but I've gone to the Gator Hardshells.

Stucky

I'm wondering if the people who complain about Gatorskins being too hash/stiff aren't inflating them too much? I was just thinking about that on today's ride- I currently have 25mm Gators- run 'em at around 100psi....and my aluminum bike (albeit, a nice-riding AL bike) rides just fine. I'd imagine iffin ya start blowin' 'em up to 120 psi, they'd be as hard as a rock.

richietables 

Either one you mentioned will be great-

Drew Eckhardt 

I rode 90-95psi front, 100-105psi rear at weights past 205 pounds plus 15 pounds of luggage.

I think it's about frame of reference.

Compared to flat resistant tires like some of Specialized's Armadillos they ride great.

Compared to supple tires they don't, even where those tires compromise for longer life and flat resistance like the GP4000S.

2 Piece

Depends on what you are looking to do with tires. 4 seasons have a fantastic ride quality, they are light weight and pretty fast. The great ride quality comes from a light weight sidewall which is very easy to get snake bite flats or is susceptible to debris cutting the sidewall. The tread area of the 4 season is very resistant to puncture types of flats, just the soft sidewall is where I had problems. But they sure did ride nice!
I have switched to Gatorskin Hardshells, a lot heavier, not near as fast on spin up, ride is a bite rough/ hard, roll very well and extremely flat resistant. But I do like them for my commute.