This is just a quick guess: Mondonico (based on the seat stay sig)
I suspect that the BB is italian (70 vs 68mm)
Aelle was good steel. Columbus' answer to 531 and about the same weight (almost). But... that sticker was probably applied after the new paint job and it is fairly questionable that it belongs there...
Measure the bottom bracket, the rear spacing, the thickness of the ST, and figure out what size seat post this takes and lots of people might help you here. Those lugs look like Cinelli to me, with a custom BB lug cut out. 99% sure that's an Italian bike.