Originally Posted by c_booth

This is OTM. And just to follow on from my last post, the steeper the headtube angle, the more upper body weight is thrown forward onto the handlebars, which itself creates a twitchier ride. It also means less fork rake is required to hit the trail sweetspot (which again will vary depending on the type of ride you're aiming for) which in turn pulls the front axle back towards the cranks - this of course shortens the wheelbase, resulting in a more responsive ride (a good thing on a track bike), but it also places your hands/weight further over the front axle, which also effects handling (to what degree I'm not so sure), and the only way to shift that weight back would be to shorten the stem, which would undoubtedly effect a further increase on the twitchiness of the ride.

So designing a frame/fork set i always think is kinda like filling in a jigsaw puzzle - sometimes the bits don't quite fit perfectly, in which case it's about compromising in a way that will least jeopardise the type of ride you're aiming to achieve.