Joined: Mar 2006
Posts: 1,916
Likes: 2
From: Perth, Australia and sometimes Penang Malaysia
Bikes: Litespeed L1r, Litespeed Ghisallo 07, TCR Advanced Team SL 0 ISP, Giant TCR Advanced SL, Giant TCR Advanced Team - T-Mobile, Giant Propel Advanced SL
It's definitely the bulbous nose on that saddle coupled with your stretched out position, why don't you just swap the toupe to the Madone and do the longer rides to check it out. I'd measure the sit bone distance to the stem so you can put the saddles in relatively the same position.
The toupe and Arione are definitely flatter designs.