When I picked up the frame from the LBS, they had a tag on the dropout stating it was set at something like 134.4mm. I took the frame home and mounted a 135mm wheel but didn't really pay attention to how it fit but I did not notice anything unusual. Over several months I took the wheels on and off several times and at one point I noticed that I had to crank the QR a lot more than usual and that the axle didn't seem to be long enough. I rode it a bit but this bothered me so I took the wheel off and took the QR skewer out and it was warped (I suspect from being used to compress the stays to fit the axle). I measured the dropouts with a digital caliper and they came out 142.2mm.

Short answer is that based on my limited recollection, the 135mm wheel did fit correctly at first.