Well, I find a brand new Brooks Swallow to be more comfortable than the C15 Cambium I have. The swallow I totally forget is there while I'm always aware of the Cambium. I'm also having good luck with the Gyes saddles. I don't know if they will last as long as a Brooks but they are a good bit cheaper and seem well made. I feel like the Gyes are a bit "springier" out of the box than the Brooks.
For comparison of the profiles
The Brooks Cambium C15:
The Brooks Swallow:
The Gyes GS-06: