In most of those nuts, the hole is off center by a differing amount, to allow the rear wheel to be aligned in the frame. Welding frames causes the metal to shrink and pull in varying degrees depending on the amount of filler metal added, fit tolerances, etc. Thru axles are not tolerant of the misalignment that welding can cause and there is no way to adjust them after fabrication. The eccentric(or off-center) nuts allow the drive side dropout to be moved forward/back/up/down as necessary to correct the alignment. This appears to be Lynskey's own version of the Syntace thru axle system, which uses similar eccentric nuts and a pinch bolt.