It can also be caused by socks that are too thick and literally keep your feet too hot. I had some older, thick cotton Descente socks that I had purchased many years ago that were still in good shape. A fitting took away most of my foot problems but they did not go completely away until my fitter suggested buying thinner, cooler socks. I purchased DeFeet Aerators and the last of my problems went away. So it literally can be temperature related.