The math part about bikes makes my head hurt.

All the other measurements for these two bikes are exactly the same. Exactly. The only difference is the 2 degrees in the headtube angle. The 74.7 is a track bike, and the 72.7 is a "road" purpose fixed gear ... which leads me to suspect that the 72.7 would have more relaxed handling ... my question I guess is will the difference (everything else being equal) be enormous? I have a road bike (a good one) and would like something with snappier handling for riding around the city on ... but, I am likely never ever going to see the inside of a velodrome.

Crap. I guess I'm just going to have to ride some bikes with similar geo. at the LBS. I hate doing that though when I know that I'm not going to buy anything from them.