The Giant was more upright, which I think made it feel more comfortable but at the same time I think would make it tougher to go fast. It also had a wider seat, which is minor, and the ride felt smoother with respect to bumps in the pavement.
Honestly the comfort factor is minor to me. The only drawback to the Trek was the price, I really liked everything else about it better, it was just more than I was looking to spend. I am leaning towards the Trek because I think the more I like the bike, the more likely I am to get on it.