Keep in mind that hub bearings come in 2 general classes, the angular contact traditional cop/cone bearings which need to be adjusted for absolutely zero play, and radial contact typical of cartridge bearings, which should be left with a bit of axial play (radial play is not adjustable).
The goal is to end up with correct adjustment for either design after the QR is tightened, so you start just slightly (very slightly) looser to allow for some compression of the hollow axle.
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