pretty sure that ones heart, lungs, training and % of muscle to body fat are biggest performance factor. Speaking as someone who used to show up at training rides with light Gucci equipment and got dusted regularly until the motor got stronger. You may perceive the difference in light wheels in an unladen bike but once you start loading it up your ability to climb a mtn at x speed will be primarily a function of total weight and not a few oz on the wheels. Calories required to climb a mtn are figured on total mass, elevation and time. If you put 10lbs on that bike the energy required to climb that mtn will be more than if you put on 4oz on the wheels, spokes, helmet or saddle. Pretty sure if you got your touring bike and put heavy touring wheels that weighed a 1/2 pound more each than light ones and road it up a mountain you'd be a hell of a lot faster than clmbing that mountain with superlight wheels and 20lbs on the bike.